Evidence for coiled-coil dimer formation by an Epstein-Barr virus transactivator that lacks a heptad repeat of leucine residues.

Flemington E; Speck SH

Division of Tumor Virology, Dana-Farber Cancer Institute, Boston, MA.

Proc Natl Acad Sci U S A 87: 9459-63 (1990)

Abstract
Two regions of the Epstein-Barr virus (EBV) BZLF1 gene product, ZEBRA, share sequence homology with c-Fos, one of which corresponds to the DNA binding domain of c-Fos. ZEBRA does not, however, contain the heptad repeat of leucines present in the dimerization domains of leucine zipper proteins. Here it is shown that ZEBRA binds its recognition sites as a homodimer and that the region adjacent to the basic DNA binding domain is essential for dimerization. This region contains a 4-3 repeat of predominantly hydrophobic residues, which is precisely in register with the hydrophobic heptad repeat present in the leucine zipper proteins with respect to the basic DNA binding domain. A mutational analysis of ZEBRA supports a model for dimerization involving a coiled-coil interaction. These results indicate that a heptad repeat of leucines is not a structural requirement for formation of coiled-coil dimers by transcription factors.

Mesh Headings

Amino Acid Sequence
DNA-Binding Proteins*
Epstein-Barr Virus*
Genes, Viral*
Leucine
Molecular Sequence Data
Mutagenesis, Site-Directed
Plasmids
Repetitive Sequences, Nucleic Acid
RNA Polymerases
Sequence Homology, Nucleic Acid
Support, U.S. Gov't, P.H.S.
Trans-Activators*
Transcription, Genetic
Translation, Genetic

Unique Identifier: 91067725

Gene Symbols

GCN4; c-JUN; c-FOS; BZLF1

Chemical Identifiers (Names)

EC 2.7.7.6 (RNA Polymerases)
(BZLF1 protein)
7005-03-0 (Leucine)